Peer-Reviewed, Original Research:

Holden, A. R. (2017) “A taphonomic and paleoenvironmental study of Late Pleistocene southern California based on a unique fossil insect assemblage from the Rancho La Brea Tar Pits.” In advanced preparation.

Holden A. R, Angus, R., Barclay, M. (2017) “Nearctic origins of a previously considered invasive species, Necrobia violecea (Coleoptera: Cleridae) to North America.” In advanced preparation.

Barclay, M., Holden, A. R. Southon, J., Angus, R. (*Author order TBD). (2017) "Chance preservation of subfossil xylophagous beetles within their associated wood substrate, provides a unique opportunity to test dating methods for wood and insect remains on known synchronous sample." In preparation.

Holden, A. R., Southon, J. R., Will, K., Kirby, M. E., Aalbu, R. L., and Markey, M.J. (2016). “A 50,000 year insect record from Rancho La Brea, southern California: Insights into past climate and fossil deposition.” Quaternary Science Reviews. Accepted.

Holden, A. R., Erwin, Diane M., Schick, K. N., and Gross, J. (2015). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.” Quaternary Research, 84(3): 358-367.

Holden, A. R., and Southon, J. R. (2015). “Radiocarbon dating and stable isotopic analysis of insect chitin from the Rancho La Brea Tar Pits, southern California.” Radiocarbon, 58(01): 99-113.

Holden, A. R., Koch, J. B., Griswold, T., Erwin, D. M., and Hall, J. (2014). “Leafcutter bee nests and pupae from the Rancho La Brea Tar Pits of southern California: Implications for understanding the paleoenvironment of the Late Pleistocene.” PLoS One, April 9.

Holden, A. R., Harris, J. M. and Timm, R. M., (2013). "Paleoecological and taphonomic implications of insect-damaged Pleistocene vertebrate remains from Rancho La Brea, southern California." PLoS One, July 3. 

Holden, A. R. and Harris, J. M. (2013).  "Late Pleistocene Coleopteran Galleries in Wood from the La Brea Tar Pits: Colonization of Juniper by Phloeosinus Chapuis (Curculionidae: Scolytinae) and Buprestidae." The Coleopterists Bulletin 67(2):155-160.

Abstracts, Non-Peer Reviewed, Popular Articles:

Southon, J., Holden, A. R. (2017) "Insects are Wood: Sample Preparation of 

Insect Cuticle for Radiocarbon Dating and Stable Isotope Measurements." Poster for the 14th Annual Conference on Accelerated Mass Spectrometry.

Presslee, S., Macphee, R. D., Collins, M., Southon, J., Holden, A. R., Farrell, A. (2016) “Radiocarbon Dating and Proteomic Analysis of Highly Purified Bone Collagen Derived from Rancho La Brea Mammal Fossils”. Poster for the Society of Vertebrate Annual Meeting.

Holden, A. R., Erwin, Diane M., Schick, K., N. and Gross, J. (2016). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.”  Poster for the American Quaternary Society 24th Biennial Meeting.

Holden, A. R., and Southon, J. (2016) “Successful methods to radiocarbon date insect chitin from the La Brea Tar Pits, Southern California.” Poster for the American Quaternary Society 24th Biennial Meeting.

Holden, A. R., (2016) Erwin, Diane M., Schick, K., N. and Gross, J. (2015). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.” Poster for the American Quaternary Association 24th Biennial Meeting.

Holden, A. R., Southon, J. (2016) “Successful methods to radiocarbon date insect chitin from the La Brea Tar Pits, Southern California.” Presentation for the American Quaternary Association 24th Biennial Meeting.

Holden, A. R., Erwin, Diane M., Schick, K., N. and Gross, J. (2015). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.”  Poster for the XXXIV Willi Hennig Society Meeting.

Holden, A. R., and Southon, J. (2015) “Successful methods to radiocarbon date insect chitin from the La Brea Tar Pits, Southern California.” Presentation for the XXXIV Willi Hennig Society Meeting.

Holden, A.R. (2015). “Nest architecture informs higher-level phylogenetic relationships within megachilid bees (Hymenoptera: Megachilidae).” Presentation for the XXXIV Willi Hennig Society Meeting.

Holden, A. R., Harris, J. M. and Timm, R. M., (2013). "Paleoecological and taphonomic implications

of insect-damaged Pleistocene vertebrate remains from Rancho La Brea, southern California." Poster

for the 73rd Annual Meeting of the Society of Vertebrate Paleontology.

Holden, A. R., Erwin, D. M., Schick, K. N., Gross, J., and Hall, J. (2013) Traces in asphalt: New

observations of Pleistocene plant – insect interactions from the Rancho La Brea Tar Pits, Los Angeles,

California, USA. Botany 2013. http://www.botanyconference.org/engine/search/index.php?func=detail&aid=887

Schick, K. N., Holden, A. R., Erwin, D. M. (2013) The Cenozoic Record of Galls: A Review. Poster for the 6th International symposium on the biology and ecology of gall inducing arthropods and related endophytes. http://6isbegia.org/images/draftprogram4july2sb2013.pdf. Queensland, Australia.

Holden, A. R., Borkent. A., and Brown, B. V. (2013).  "Progress of the Costa Rican ZADBI Project."  Fly Times, Issue 50.