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Peer-Reviewed, Original Research:

Holden, A. R. (2017) “A taphonomic and paleoenvironmental study of Late Pleistocene southern California based on a unique fossil insect assemblage from the Rancho La Brea Tar Pits.” In advanced preparation.


Holden A. R, Angus, R., Barclay, M. (2017) “Nearctic origins of a previously considered invasive species, Necrobia violecea (Coleoptera: Cleridae) to North America.” In advanced preparation.


Barclay, M., Holden, A. R. Southon, J., Angus, R. (*Author order TBD). (2017) "Chance preservation of subfossil xylophagous beetles within their associated wood substrate, provides a unique opportunity to test dating methods for wood and insect remains on known synchronous sample." In preparation.


Holden, A. R., Southon, J. R., Will, K., Kirby, M. E., Aalbu, R. L., and Markey, M.J. (2016). “A 50,000 year insect record from Rancho La Brea, southern California: Insights into past climate and fossil deposition.” Quaternary Science Reviews. Accepted.


Holden, A. R., Erwin, Diane M., Schick, K. N., and Gross, J. (2015). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.” Quaternary Research, 84(3): 358-367.


Holden, A. R., and Southon, J. R. (2015). “Radiocarbon dating and stable isotopic analysis of insect chitin from the Rancho La Brea Tar Pits, southern California.” Radiocarbon, 58(01): 99-113.


Holden, A. R., Koch, J. B., Griswold, T., Erwin, D. M., and Hall, J. (2014). “Leafcutter bee nests and pupae from the Rancho La Brea Tar Pits of southern California: Implications for understanding the paleoenvironment of the Late Pleistocene.” PLoS One, April 9.


Holden, A. R., Harris, J. M. and Timm, R. M., (2013). "Paleoecological and taphonomic implications of insect-damaged Pleistocene vertebrate remains from Rancho La Brea, southern California." PLoS One, July 3. 


Holden, A. R. and Harris, J. M. (2013).  "Late Pleistocene Coleopteran Galleries in Wood from the La Brea Tar Pits: Colonization of Juniper by Phloeosinus Chapuis (Curculionidae: Scolytinae) and Buprestidae." The Coleopterists Bulletin 67(2):155-160.


Abstracts, Non-Peer Reviewed, Popular Articles:

Southon, J., Holden, A. R. (2017) "Insects are Wood: Sample Preparation of 

Insect Cuticle for Radiocarbon Dating and Stable Isotope Measurements." Poster for the 14th Annual Conference on Accelerated Mass Spectrometry.


Presslee, S., Macphee, R. D., Collins, M., Southon, J., Holden, A. R., Farrell, A. (2016) “Radiocarbon Dating and Proteomic Analysis of Highly Purified Bone Collagen Derived from Rancho La Brea Mammal Fossils”. Poster for the Society of Vertebrate Annual Meeting.


Holden, A. R., Erwin, Diane M., Schick, K., N. and Gross, J. (2016). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.”  Poster for the American Quaternary Society 24th Biennial Meeting.


Holden, A. R., and Southon, J. (2016) “Successful methods to radiocarbon date insect chitin from the La Brea Tar Pits, Southern California.” Poster for the American Quaternary Society 24th Biennial Meeting.


Holden, A. R., (2016) Erwin, Diane M., Schick, K., N. and Gross, J. (2015). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.” Poster for the American Quaternary Association 24th Biennial Meeting.


Holden, A. R., Southon, J. (2016) “Successful methods to radiocarbon date insect chitin from the La Brea Tar Pits, Southern California.” Presentation for the American Quaternary Association 24th Biennial Meeting.


Holden, A. R., Erwin, Diane M., Schick, K., N. and Gross, J. (2015). “Late Pleistocene galls from the La Brea Tar Pits; Implications for cynipine wasp and native plant distribution in southern California.”  Poster for the XXXIV Willi Hennig Society Meeting.


Holden, A. R., and Southon, J. (2015) “Successful methods to radiocarbon date insect chitin from the La Brea Tar Pits, Southern California.” Presentation for the XXXIV Willi Hennig Society Meeting.


Holden, A.R. (2015). “Nest architecture informs higher-level phylogenetic relationships within megachilid bees (Hymenoptera: Megachilidae).” Presentation for the XXXIV Willi Hennig Society Meeting.


Holden, A. R., Harris, J. M. and Timm, R. M., (2013). "Paleoecological and taphonomic implications

of insect-damaged Pleistocene vertebrate remains from Rancho La Brea, southern California." Poster

for the 73rd Annual Meeting of the Society of Vertebrate Paleontology.


Holden, A. R., Erwin, D. M., Schick, K. N., Gross, J., and Hall, J. (2013) Traces in asphalt: New

observations of Pleistocene plant – insect interactions from the Rancho La Brea Tar Pits, Los Angeles,

California, USA. Botany 2013. http://www.botanyconference.org/engine/search/index.php?func=detail&aid=887


Schick, K. N., Holden, A. R., Erwin, D. M. (2013) The Cenozoic Record of Galls: A Review. Poster for the 6th International symposium on the biology and ecology of gall inducing arthropods and related endophytes. http://6isbegia.org/images/draftprogram4july2sb2013.pdf. Queensland, Australia.


Holden, A. R., Borkent. A., and Brown, B. V. (2013).  "Progress of the Costa Rican ZADBI Project."  Fly Times, Issue 50.

In the News:

Richmond Public Radio, 88.9FM, "What's Bugging You":

Interview with Dr. Arthur V. Evans, and producer, Steve Clark, for "What's Bugging You", a weekly radio segment on insects, which airs during "Morning Addition".

“Anna Holden, a doctoral student at the Richard Gilder Graduate School at the American Museum of Natural History in New York, shares her insect research at the La Brea Tar Pits in California."

http://ideastations.org/radio/news/insects-la-brea-tar-pits


Science Magazine:

Drowning in tar, eaten by insects.

http://news.sciencemag.org/2013/07/scienceshot-drowning-tar-eaten-insects

Ancient bees pulled from tar pit.

http://news.sciencemag.org/plants-animals/2014/04/scienceshot-ancient-bees-pulled-tar-pit


Smithsonian Magazine:

Ice-Age bees uncovered at the La Brea Tar Pits.

http://www.smithsonianmag.com/smart-news/ice-aged-bees-uncovered-la-brea-tar-pits-180950507/?no-ist


National Geographic:

Tar pit bees connect California’s past to the present.

http://phenomena.nationalgeographic.com/2014/04/12/tar-pits-bees-connect-californias-past-to-the-present/


NBC:

Bee fossils provide rare glimpse into Ice Age environment.

http://www.nbcnews.com/id/54923183/ns/technology_and_science-science/t/bee-fossils-provide-rare-glimpse-ice-age-environment/#.VDXmXtylnFI


NPR Science Friday:

Prehistoric leafcutter bee pupae.

http://www.sciencefriday.com/articles/insect-microfossils-provide-prehistoric-insights/

Tar Noir: Paleoforensics at the La Brea Tar Pits.

http://www.sciencefriday.com/video/08/21/2014/tar-noir-paleoforensics-at-the-la-brea-tar-pits.html


Science News:

La Brea Tar Pits yield exquisite Ice Age bees.

https://www.sciencenews.org/article/la-brea-tar-pits-yield-exquisite-ice-age-bees


Science Daily:

Fossil insect traces reveal ancient climate, entrapment, and fossilization at La Brea Tar Pits.

http://www.sciencedaily.com/releases/2013/07/130703101433.htm


Science News Daily:

Fossil insect traces reveal ancient climate, entrapment, and fossilization at La Brea Tar Pits.

http://www.sciencenewsdaily.org/archaeology-fossils-news/cluster388843272/


Science Magazine News for Kids:

Tar pit bones yield climate clues: Insect damage to ancient bones reveals length of ‘summer’

during the last Ice Age.

http://www.sciencenewsforkids.org/2013/07/insect-damage-to-ancient-bones-reveals-lengthof-

summer-during-the-last-ice-age/


FOX News:

Bee fossils provide rare glimpse into Ice Age environment.

http://www.foxnews.com/science/2014/04/11/bee-fossils-provide-rare-glimpse-into-ice-age-environment/


Los Angeles Times:

A web search in the Santa Monica Mountains.

http://articles.latimes.com/2010/apr/18/local/la-me-spiders18-2010apr18

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